Tsukasa, a college student, is rescued from an attack by a devil, one of many vampires that can blend in among the human population. Anzai, her savior, is a half-devil who exploits his supernatural gifts as a member of a shadowy police task force that specializes in devil-related crime in Tokyo. As Anzai continues to keep guard over Tsukasa, the two quickly forge a tentative bond—one that Anzai fears will test his iron-clad rule of never drinking human blood...
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Miyako Arata is newly assigned to the Shinjuku Ward Office's Nighttime Regional Relations Department. Each of Tokyo's 23 wards has one such department, established to mitigate paranormal and occult-related events. Arata's special skill is the understanding of non-human speech, and the story begins with him encountering a youkai at Shinjuku Gyoen park who refers to him as the legendary Heian-era exorcist, Abe no Seimei.
Narumi is certain that Masaru will be kidnapped by wooden puppets with supernatural strength after fighting them with all his strength. Masaru inherited his father's wealth which has brought many enemies out of the shadows, too many for Narumi to defend him alone. When Narumi is feeling discouraged, Masaru's watcher, Shirogane, arrives with a powerful weapon, the puppet Arlequin.
In order to survive the eventual arrival of the "Dusk End," people of the western reaches of the "Land of Dusk" devoted their efforts to rediscover and recreate lost alchemic technologies. Those technologies were then gathered in the alchemy city known as "Central," where research was conducted on how to halt the advance of the twilight. Having learned the newest alchemic techniques in Central, young alchemist Logy requested a transfer to an understaffed town of Colseit on the frontier to make use of his abilities. There he meets Escha, an apprentice girl assigned to the Development Department. Together, the two make a promise to use their alchemic techniques to bring success to the Development Department.
When interdimensional travelers from an otherworldly land known as "Japan" appear, death and destruction always follow. It's up to Executioners like Menou to exterminate the Lost Ones before they wreak havoc. When Menou encounters a Lost One named Akari, it's bloody business as usual... until Menou discovers Akari can cheat death. Even so, Menou has a job to do, and she is committed to her executioner's mission come hell or high water — provided her newly stirring feelings don't get in the way.
After saving humanity from the despicable Demon King, Leo Demonhart does not earn a hero's standing. Instead, he is regarded with suspicion and hostility by those he fought to protect. Treated as a pariah with nowhere else to turn, exiled Leo seeks a position in the army of his former nemesis! But the army is not what it once was before its defeat at Leo's hands, so to maintain his new position, Leo must whip the forces back into fighting shape.
Following the success in the Camelot Singularity, Ritsuka Fujimaru and Mash Kyrielight are assigned to the last Singularity in the Grand Order initiative. In Ancient Babylonia, 2500 BCE, they embark on a mission to secure humanity's survival. Upon arrival, they learn that three gods have threatened Uruk, the Babylonian city ruled by King Gilgamesh. Ritsuka and Mash must work together to fend off the invasion of mysterious beasts in Uruk under Gilgamesh's orders while investigating the true nature of the three gods' actions against humanity; but unknown to Ritsuka, an ancient entity is slowly rising from its slumber.