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Nick and his mother team up to fight the Mauvais Dentes. Meanwhile, details about Juliette's affliction come to light as Monroe and Rosalee race against the clock to save her; Nick gets into trouble with the FBI; and Capt. Renard deals with his brother.
The fourth season ends with a nuclear explosion in the Pacific. The event reignites a search for stolen nuclear weapons, and leads Hetty to shuffle partnerships. She sends Callen and Kensi overseas and leaves Sam and Deeks as a stateside team.