The ongoing saga of a well-to-do but down-to-earth California clan of diverse personalities, from a liberal gay lawyer to a conservative talk-show host, and how their lives change after the death of a patriarch reveals that the family business is in trouble.
Nora decides to follow her heart with Brody, though it's not a popular decision among her kids; Sarah and Luc mull adding to their family; Scotty's chance encounter with his and Kevin's former surrogate (Roxy Olin) makes him wonder where's she's been.
Under the guise of surprising Evan on his birthday, Nora and Sarah visit Washington, D.C., but they really want to find out what Kitty's been up to on the East Coast. Meanwhile, Kevin and Scotty finally become Olivia's parents, but their new daughter is a tad uneasy about starting over.
Valentine's Day celebrations go awry when Tommy tells Sarah about another secret that Nora has been keeping hidden for years, while Saul's night out with Jonathan (Richard Chamberlain) is ruined by a well-known restaurant critic.
Nora and Holly's reminiscing triggers a revelation about Nora's past that could have a major effect on Holly's future in California. Meanwhile, Justin is annoyed by the reappearance of Annie's former boyfriend; Kevin and Scotty take another look at their adoption prospects; and Saul tries to summon forgiveness for Jonathan.
Nora's otherwise harmless date night somehow morphs into a scandal for the other Walker couples. Meanwhile, Kitty's relationship with Seth attracts unwanted attention on the Internet and from her boss.
Justin finds another reason to continue to visit Holly; Nora is impressed by her radio colleague's handling of Sarah and Kitty's age-old quarrels; Paige helps Kevin and Scotty patch up their relationship woes.
When Kevin's relationship with Scotty is rocked by the latter's infidelity, the entire family is drawn into a dramatic confrontation. Meanwhile, Kitty is at odds with Jack's latest pursuits; and Sarah and Luc unveil surprising news.
Nora's radio days may be over before they started when an audition pits her against a celebrity doctor (Debra Monk); Kitty decides to jump back into the dating pool with her handyman (Jeremy Davidson) in Ojai.
Holly's memory lapses take a toll on her psyche; Kitty and Saul do some soul searching over their romantic dilemmas; Justin and Rebecca wonder where their relationship is headed; Cooper gets a lead role in a production of "Romeo and Juliet."
Still hiding out in Ojai on her journey of self-discovery, Kitty may finally be ready to move on when she meets a handsome handyman (Jeremy Davidson). Meanwhile, the Walkers secretly plan a birthday party for Sarah, though it's Luc who could be surprised by her actual age.
Luc's career as an underwear model takes off at a launch party, where teenage Paige engages in some questionable behavior. Meanwhile, Nora's part-time job as a florist may come to an end after an offer from a radio-show producer; Kevin deals with a troubled runaway; and Justin tries to help Holly jog her memory.
The fifth season begins one year after the Walkers were shaken by a horrific car accident, and now the somewhat estranged family prepares for an uneasy reunion on the occasion of a homecoming party for Justin. Meanwhile, Kevin is practicing law again; Sarah is obsessed with Narrow Lake; Saul and Scotty's restaurant is up and running; and no one wants to discuss Kitty's personal life.