Betty won Daniel and Wilhelmina's contest and became a features editor at Mode. She broke up with Matt, who saw her kiss Henry and then, out of spite, decided to become her boss.
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Daniel Meade (Eric Mabius)
Newlywed Daniel tended to his ailing bride, but to no avail: She succumbed to cancer in the Season 3 finale.
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Wilhelmina Slater (Vanessa Williams)
Willy had a rough year. A DNA test determined that William was not her biological child. Her boyfriend fled after bilking Meade for millions. And she was dethroned as Mode's editor-in-chief by Claire Meade.
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Marc St. James (Michael Urie)
Marc lost the coveted editor's position to Betty. He's still Willy's trusted assistant, but he's more revenge-minded than ever.
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Amanda Tanen Sommers (Becki Newton)
Amanda's loyalties are split when Marc and Betty compete for an editorial position, but all that talk about career advancement has her thinking as well.
6 of 8
Ignacio Suarez (Tony Plana)
Ignacio was on the mend after suffering a heart attack and losing his girlfriend, who moved out of town for a nursing job.
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Hilda Suarez (Ana Ortiz)
Hilda's relationship with Archie is going well, but she's starting to get cold feet about moving forward with the upstanding city councilman.
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Justin Suarez (Mark Indelicato)
Justin's dreams were dashed when he was not accepted at the performing-arts high school. Instead, he'll enroll in public school in the fall, which promises to be a challenge for the stylish teen.