In the third-season premiere, Betty is finally ready to make a life-altering decision after fielding proposals from Gio and Henry. Meanwhile, Ignacio gets a new job at a fast-food restaurant, and Daniel deals with the fallout of Wilhelmina's power play. Lindsay Lohan guest stars as Kimberly, Betty's high-school nemesis. Regis Philbin and Kelly Ripa appear as themselves.
New Mode editor Wilhelmina tries to entice Betty to take a job as her assistant; Daniel's desire to keep his son in the U.S. hits a snag in the form of a custody battle; Christina finds reason to be wary of her estranged husband's motives.
Betty does her best to tell a police detective (Mark Consuelos) what happened in an accident that shocks the folks at Mode. Meanwhile, Ignacio learns some surprising information about Hilda and Coach Diaz.
Ignacio flexes his new citizenship muscles by getting ready to vote for the first time, while the upcoming election brings Hilda a new customer (Ralph Macchio). Also: Betty's rivalry with Kimmie (Lindsay Lohan) resumes, and Wilhelmina finds yet another way to get under Daniel's skin. Supermodel Adriana Lima has a cameo as a Mode cover girl.
Betty tries to nip a potential public-relations catastrophe in the bud by halting the distribution of a new issue of Mode, but it's made difficult by the fact that Daniel is incommunicado at a company retreat. Meanwhile, Hilda's salon permit is rejected by the city, and Connor's fiancée meets the magazine's staff.
Betty finds herself in competition with Marc when both apply to a renowned training program for young editors. Meanwhile, Justin strikes up an unlikely friendship; Daniel is smitten with a new woman; and Wilhelmina tries to get Grant's attention. Designers Mark Badgley and James Mischka appear as themselves.
Betty fights an uphill battle to make a good impression on a YETI editor (Bernadette Peters), and meets her counterpart (Nikki Blonsky) at Elle magazine as the publication's rivalry with Mode continues. Elle editors Robbie Myers and Joe Zee appear as themselves.
Betty deals with a multitude of distractions, including an apparently renewed interest from her neighbor, as she mulls an important decision that will affect her family and bank account. Meanwhile, Connor vows to identify the mystery man in Molly's life, which sends Daniel running for cover.
Betty's attempt to create a video tribute to Claire on her 60th birthday winds up shining an accidental spotlight on Ignacio. Meanwhile, Daniel and Wilhelmina face similar obstacles in their new relationships.
Fashion Week in New York City makes Betty's life more hectic than usual as she helps an eccentric designer put on a show and partners up for a YETI assignment with someone who works at a sports magazine. Meanwhile, Wilhelmina worries about her new relationship's effect on her career, and Daniel hopes to see another side of Molly.
A YETI assignment grants a shocking look at Mode's financial history for Marc and Betty, who also copes with news that Henry has posted on the Internet and deals with increased attention from a previous YETI partner, Matt (Daniel Eric Gold).
Betty meets Matt's mother (Christine Baranski); Hilda's family try to set her up with Archie (Ralph Macchio); Wilhelmina is faced with financial difficulties; Daniel's relationship with Molly undergoes another adjustment.
Marc and Amanda give Betty a makeover to take her romance with Matt in a new direction; Daniel and Wilhelmina try to spice up the next issue to improve Mode's bottom line; Hilda ponders her relationship with Archie (Ralph Macchio).
When Betty and her family are on the guest list for a renowned Easter egg hunt thrown by Matt's obscenely wealthy father (David Rasche), Daniel hatches an idea that could improve Mode's financial situation. Meanwhile, Wilhelmina finds a reason to be suspicious of her son's DNA.
The hope that Matt's wealthy father will save Mode from financial collapse is threatened by the kidnapping of an heir to the Meade empire. Meanwhile, Archie Rodriguez (Ralph Macchio) tries to enlist Hilda's help with his political aspirations.
Matt's behavior is a cause for concern when he works with Marc and Betty on a photo shoot with Grammy-winning singer Adele for their final YETI assignment. Meanwhile, Betty fields a request from Matt's mother (Christine Baranski); Claire and Wilhelmina joust over their positions at the magazine; and the Mode universe witnesses a pair of wedding proposals.
Betty's plans to move in with Matt are interrupted by a surprise visit from Henry (Christopher Gorham); Wilhelmina uses the Hartley family's dysfunction to renew her rivalry with Claire; Daniel's new lifestyle makes him the toast of the town. Cameos include Billie Jean King, Elisabeth Hasselbeck and Joy Behar.
Betty and Marc are candidates for a job opening at Mode after an editor's sudden death; Wilhelmina tries blackmail to get Claire out of the picture once and for all; Justin hopes to be accepted to a performing-arts school. Cameos include Rachel Maddow and Robert Verdi.