Henry comes back to his old job at Mode, but Betty is distracted by the dilemma presented by a tempting offer from Wilhelmina and her loyalty to Daniel. Meanwhile, Bradford uses Alexis' amnesia to his advantage, Marc gets the most out of Justin's internship and Hilda takes one last trip down memory lane.
A new sandwich vendor (Freddy Rodriguez) at the office triggers ambivalence in Betty. Meanwhile, the Mode staffers are all atwitter over the Meade's annual Black and White Ball, where Wilhelmina hopes to put her wedding plans back on track, and Amanda gets ready to announce her famous bloodlines.
Alexis' return to work is complicated by some unladylike behavior; Daniel is troubled by demands from a major advertiser (James Van Der Beek); Betty enrolls in a writing class; Amanda begins the search for her father.
Henry is upset by Betty's foray into online dating, which results in a rendezvous at a bowling alley; Daniel and Alexis prepare for dinner with Wilhelmina; Hilda is oblivious to Justin's new friends; Marc is smitten with a photographer.
As her wedding approaches, Wilhelmina worries about being overshadowed by her maid of honor, Victoria Beckham. Meanwhile, Ignacio prepares to become a U.S. citizen, but his joy is tempered by Betty's decision to move out of the house---and in with Henry.
With some assistance from Mode's weekend receptionist (Mo'Nique), Henry works overtime to help Betty get something she needs; Wilhelmina mulls a new plan of attack in the wake of the latest episode in her romantic saga with Bradford.
When a crisis caused by Wilhelmina's sabotage forces the Mode staff to redo an entire issue, Betty and Daniel try to spring a cover girl (Eliza Dushku) from rehab for another photo shoot, while Alexis attempts to coax more time out of the magazine's printer.
Amanda seeks help from a clairvoyant (Annie Potts) in the search for her father's identity; a visit to the magazine by Justin and his classmates prompts Betty to coax Daniel to change the image of Mode's models; a new twist develops in Wilhelmina's designs on the Meade empire. Rappers Bow Wow and Omarion appear as themselves.
Betty's new perfume---a fragrance dreamed up by the deceased Fey Sommers---has bizarre effects on herself and Henry; Claire goes on trial for killing Fey; Wilhelmina continues her quest to be part of the Meade family.
Betty's assignment to interview a writer is spoiled by a case of mistaken identity; Daniel is surprised to learn he's closer to his new love interest (Gabrielle Union) than he imagined; Marc helps Amanda in her quest to meet her biological father.
Gio may have a solution to Betty's problems when her plans to spend her 24th birthday with Henry are threatened by Charlie's surprise visit. Meanwhile, Wilhelmina focuses her wicked ways on Daniel's relationship with her sister, Renee (Gabrielle Union).
Renee's delusions have her convinced that Betty is in love with Daniel. Meanwhile, Hilda's latest client is her old nemesis, Gina Gambarro, who comes back to Queens to flaunt her new, wealthy lifestyle as a doctor's wife. "Project Runway" winner Christian Siriano and judge Nina Garcia have cameos.
Betty tries a different tactic in her rivalry with Charlie by throwing her the mother of all baby showers. Meanwhile, Justin's poor grades in gym prompt a confrontation between Hilda and his teacher (Eddie Cibrian); Daniel is sent reeling by relationship woes; and Christina learns the identity of the father of the child she's carrying for Wilhelmina.
Betty volunteers to chaperone Justin's middle-school dance, only to find that Gio has made the same commitment. Meanwhile, Amanda prepares for a reality show with her father, KISS bassist Gene Simmons; Hilda does her best to get the attention of Coach Diaz; and Wilhelmina returns to Mode.
In the second-season finale, Betty weighs the pros and cons of a sudden romantic dilemma. Meanwhile, Mode faces supermodel Naomi Campbell and the staff of Elle magazine in a charity softball game, Hilda hears surprising information about Coach Diaz, and a young French boy makes a claim that shocks the Meade empire. Kimberly: Lindsay Lohan.