Betty scrambles to find a missing mock-up of the magazine before its less-than-flattering pictures of a movie star (Sarah Jones) fall into the wrong hands. Meanwhile, Bradford takes a personal interest in the Fey Sommers affair.
Crises make for strange bedfellows when Betty, Marc and Amanda try to find out how a secret magazine feature was scuttled, and Wilhelmina and Daniel become allies to beat a deadline on a photo spread. Meanwhile, Justin spends a day at Mode working on a school project.
Betty deals with the embarrassment of her Halloween costume, the nervousness of a new crush and has her hands full finding Daniel's missing watch. Meanwhile, Daniel visits his mother (Judith Light), and Wilhelmina comes apart at the seams over a dress size.
Executive producer Salma Hayek guest stars as a woman who makes an immediate impression on playboy Daniel. Meanwhile, Wilhelmina gets a surprise visit from her daughter, and Betty and Hilda face a financial crisis.
Betty is asked to review a fancy hotel; Wilhelmina covets advertising dollars from a wealthy Texan (Brett Cullen); and Sofia (Salma Hayek) and her entourage throw their weight around, much to Daniel's chagrin.
Martha Stewart has a cameo as the Mode staff marks Thanksgiving in different ways, including Daniel's ulterior motive for attending his parents' brunch and Wilhelmina's first attempt at cooking a turkey.
The day after Thanksgiving turns into a turkey for Betty, who must hastily organize a celebrity photo shoot, and Daniel, who is reeling from a hangover and a snub by Sofia. Meanwhile, Wilhelmina and Marc wind up on the wrong side of the tracks.
Amanda hopes her successful holiday office party will be the gift that keeps on giving. Meanwhile, Betty sorts through her feelings for Walter and Henry; Daniel's love for Sofia grows; and Marc tries to stay on Wilhelmina's good side.
Christina's swag lottery creates a feeding frenzy among the office fashionistas, while Betty wears herself out helping a suddenly cash-poor Daniel entertain a famous Japanese designer (Kurando Mitsutake).
Daniel's future plans with Sofia are thrown into question when Betty makes a startling discovery; Wilhelmina's romance with Ted changes her mood at the office; and Ignacio doesn't see eye-to-eye with his immigration caseworker.
Betty helps Daniel mend his broken heart by urging him to return to what he knows best; the masked woman is revealed; Wilhelmina makes personnel decisions for her hypothetical ascension at Mode; Hilda scrambles to find a new source of income.
Betty has her work cut out for her getting ready for Mode's big event during New York City's Fashion Week; Marc and Amanda can't wait to get their hands on something everybody wants; Christina hopes to make her mark in the design world; and Daniel asks Hilda to give her sister a hand. Katharine McPhee has a cameo as herself.
Betty finds herself in a romantic dilemma while being forced to play peacemaker during the post-Fashion Week chaos at Mode. Meanwhile, Justin gets in trouble at school, but Hilda bristles at Santos' reappearance in her son's life.
Daniel deals with a family matter and turns to former classmate Grace Chin (Lucy Liu) for legal help; Alexis meets a guy (Jerry O'Connell) while stranded at a sports bar with Wilhelmina during a blizzard; and Betty makes a new friend with a connection to Henry.
Betty's orthodontist becomes her date for Charlie's birthday party; Wilhelmina tries to make Daniel's hiring of Grace Chin for his father's defense come between the Meade siblings; and Amanda attracts the attention of a designer.
Betty reluctantly returns a favor from Marc by agreeing to pose as his significant other during a visit from his meddlesome mother (Patti LuPone). Meanwhile, an executive order puts a strain on Alexis and Daniel's working relationship.
Betty works overtime when Daniel finds old habits hard to break; a gossip scribe with Fey Sommers' diaries creates opportunities for Alexis and Wilhelmina; and Ignacio struggles to maintain a professional relationship with the amorous Constance.
Betty tries to distance herself from a string of recent disappointments; Alexis fields her first offer for a date since her operation; Santos surprises Hilda; Daniel deals with the fallout from a one-night stand; and Amanda makes assumptions about a current suitor.
Betty struggles to find money to solve another Suarez family dilemma while planning Mode's Assistants Week bash at a medieval-themed restaurant with an unlikely tie to Amanda's past. Meanwhile, Daniel tries to heed Betty's advice about his sexual-addiction problem; Alexis considers Rodrigo's offer; Hilda faces a tough cosmetology-school exam; and Wilhelmina becomes even bolder in her plotting for total Mode domination. Rodrigo: Cristian de la Fuente.
Ignacio's immigration woes take the family to Guadalajara, where Betty asks an aunt (Rita Moreno) about her mother; Alexis runs into her last girlfriend before she had her operation; Marc feels cast aside as Amanda's best friend.
Daniel's debauchery continues unabated, and Wilhelmina reaps the benefits of her ambition. Guest stars include Kristin Chenoweth and Angélica Vale, who stars in the popular Mexican telenovela “La Fea Más Bella.”