Capt. Sharon Raydor (Mary McDonnell) leads the L.A.P.D.'s Major Crimes division in this spin-off from "The Closer," featuring characters from that series. In the premiere, the newly appointed Raydor investigates a string of store robberies that turns fatal. Meanwhile, an undercover detective proves her worth and is transferred into the division; and Raydor tries to protect Rusty.
As Raydor works to gain respect in the division, the Major Crimes crew works to crack the case of a brutal and complicated homicide. Meanwhile, Raydor tries to supervise Rusty; and Provenza struggles to accept all the changes in the department.
The unit joins forces with the FBI when a homicide case spins around the Israeli mob and an "intuitive life strategist." Elsewhere, Provenza displays his vulnerable side, and Rusty's first school day takes an unexpected turn.
The aftermath of a deadly shooting on gang turf finds Raydor making a daring maneuver that could threaten her career and jeopardize Det. Sykes' life. Elsewhere, Rusty makes it quite apparent that he has no interest in meeting his biological father.