Michelle's shooting causes problems and turmoil for Alex, and his family try to help him. Meanwhile, Victor and Natalie want to push up their escape; and Oscar poses as a Boston native when he gets acquainted with his new New York City neighbor.
Natalie relies on an outsider to obtain crucial info for her. Meanwhile, Sam and Alex make a dangerous move to detain an SVR source; and Katya implores Alex to give her some breathing room with the task force, which leads to Alex meeting Victor.
The FBI think they have a mole, so an investigation gets underway, and it puts Alex at war with his own ethics. Elsewhere, Victor devises a new method for staying on top of the O'Connors; and Natalie makes a grievous error while searching for a lead.
A surprising discovery leaves Alex reeling as he tries to adjust his theories in light of what he found at Philadelphia City Hall. Meanwhile, the Rezident questions Katya and Mark's dedication; and Natalie discovers the dangers of Black Dagger.
Alex and the FBI head to Philadelphia, where they search for Mikhail's hidden laptop, but Victor instructs the O'Connors to snatch the computer first. Meanwhile, Katya uncovers something Natalie's been hiding and pressures her to make a difficult choice.
Katya manipulates Alex and makes him think he's mistaken when he almost exposes his parents as spies, but her lies lead to a rift in the family. Meanwhile, Victor is warned to keep the O'Connors under control; and Alex teams up with an FBI agent.