Milano will join returning judges Isaac Mizrahi and Georgina Chapman for the Lifetime show's spin-off of Project Runway, which originated on Bravo. Milano replaces Season 2 host Carolyn Murphy. Angela Lindvall hosted Season 1.
Lifetime has yet to announce the designers competing this season, but production is slated to start later this month.
Milano currently stars in the ABC series Mistresses, which debuts tonight at 10/9c.