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The doctors receive bad news from Jackson. Meanwhile, Amelia arrives in Seattle for an unexpected visit; Richard heads to Boston to surprise Catherine; and Bailey makes a bold decision after growing disappointed with the lack of progress in her research.
Cristina seeks guidance from Meredith as she makes a decision about her future. Meanwhile, Derek and Amelia join forces to operate on conjoined twins; Jo takes on too many cases; and Stephanie uncovers the truth about Bailey's bubble-boy case.
Amelia opens up to Meredith about her time in Seattle. Elsewhere, bad news is delivered to Callie; Bailey faces the consequences of her actions; and the residents are alarmed to hear that one of them will soon be out of a job.